Tj -69 -12 TD 0.1096 Tc -0.1096 Tw (The notation +/) Tj 56.25 0 TD 0.003 Tc 0 Tw (-) Tj 3 0 TD -0.0053 Tc 0.0887 Tw ( means positive or negative effects on fitness at ) Tj -59.25 -12 TD -0.1298 Tc 0.1298 Tw (the cell or organism level.) Okasha’s book makes the sort of contribution that simply will not be able to be ignored by anyone interested in this field for many years to come. Conflict leads) Tj 229.5 0 TD 0 Tc 0 Tw (\227) Tj -229.5 -11.25 TD 0.1899 Tc 2.3012 Tw (through the evolution of adaptations that reduce it) Tj 221.25 0 TD 0 Tc 0 Tw (\227) Tj 9.75 0 TD 0.3322 Tc 0.2303 Tw (to ) Tj -231 -11.25 TD 0.0631 Tc -0.2506 Tw (greater individuality and harmony for the organism.) Fitness ) Tj -204 -11.25 TD 0.1287 Tc 0.0588 Tw (variation at the cell and group levels translates directly into ) Tj 0 -11.25 TD 0.1762 Tc 2.8506 Tw (the opportunity for within and between group change.) Evolution by Natural Selection 2. These statements best describe the concept of A. spontaneous mutation B. reproductive isolation C. homeostasis D. evolution 5. %PDF-1.3 %���� We have been ) Tj -41.25 -11.25 TD 0.1384 Tc -0.0759 Tw (especially interested in the features of organisms that affect ) Tj 0 -11.25 TD 0.1702 Tc 0.8923 Tw (the opportunity for within and between group change for ) Tj T* 0.1295 Tc 1.033 Tw (they shape the variation and heritability at the new level. ) Multi-Level Selection and the Major Transitions in Evolution The body of ideas known loosely as ‘multi-level selection theory’ takes as its 3. Tj 47.25 0 TD /F2 9.75 Tf 0.0899 Tc -0.2774 Tw (Proc. Tj 15.75 0 TD 0 Tc -0.1875 Tw ( ) Tj -139.5 -11.25 TD -0.2813 Tc 0.0938 Tw (3. ) A two ) Tj T* 0.5411 Tc 0 Tw (sp) Tj 9.75 0 TD 0.0823 Tc 3.1588 Tw (ecies group formed by association of members from) Tj 0 Tc -0.9375 Tw ( ) Tj -9.75 -11.25 TD 0.1754 Tc -0.1962 Tw (different species is indicated by the larger touching shapes; ) Tj 0 -11.25 TD 0.1481 Tc 2.7582 Tw (the size of each touching shape indicating the relative) Tj 0 Tc -0.1875 Tw ( ) Tj T* 0.2006 Tc 3.0807 Tw (proportions of each species within the group and the) Tj 0 Tc -0.1875 Tw ( ) Tj T* 0.2049 Tc 0.3576 Tw (shading of each shape indicating th) Tj 146.25 0 TD 0.0902 Tc 0.0973 Tw (e relative proportion of ) Tj -146.25 -11.25 TD 0.1959 Tc -0.1584 Tw (the two phenotypes within each species in the group. Nat.) Tj 152.25 0 TD 0 Tc -0.1875 Tw ( ) Tj -152.25 -11.25 TD ( ) Tj 9.75 0 TD 0.2019 Tc -0.3894 Tw (What happens during an evolutionary transition to a new ) Tj -9.75 -11.25 TD 0.2231 Tc 0 Tw (higher) Tj 25.5 0 TD 0.0949 Tc 5.6239 Tw ( level unit of individuality, in this case the) Tj 0 Tc -0.1875 Tw ( ) Tj -25.5 -11.25 TD 0.0809 Tc -0.0541 Tw (multicellular organism? We have studied two different kinds of ) Tj 0 -11.25 TD 0.1926 Tc 3.9699 Tw (cooperation, depending upon whether the cooperative) Tj 0 Tc -0.1875 Tw ( ) Tj T* 0.1061 Tc 0.3883 Tw (phenotype is costly or beneficial at the cell level. Mechanisms of human sexual selection Sexual selection can operate through multiple mecha-nisms, including contests, mate choice, scrambles, sexual coercion, and sperm competition (Table 1). Within each species ) Tj T* 0.1636 Tc 0.2322 Tw (there are two phenotypes \(cooperate or defect, cultivate or ) Tj T* 0.1766 Tc 0.5359 Tw (exploit, etc.\) indicated by open and shaded shapes. new work on the levels of selection, some of which has significantly re-defined the terms of the traditional debate. Tj 108 0 TD -0.0633 Tc 1.0008 Tw ( II. On this approach, the level of entity actively selected (the interactor) benefits from evolution by selection at that level through its acquisition of adaptations. The subscript ) Tj T* /F2 9 Tf -0.252 Tc 0 Tw (j) Tj 2.25 0 TD /F1 9 Tf 0.0011 Tc -0.0011 Tw ( refers to the number of non) Tj 100.5 0 TD 0.003 Tc 0 Tw (-) Tj 3 0 TD 0.0083 Tc -0.0083 Tw (mutant cells in the offspring group;) Tj 126.75 0 TD 0 Tc 0 Tw ( ) Tj -232.5 -12 TD /F2 9 Tf -0.252 Tc (j) Tj 2.25 0 TD /F1 9 Tf 0.0174 Tc 2.2326 Tw ( = 0, 1, 2, ...) Tj 54.75 0 TD /F2 9 Tf -0.003 Tc 0 Tw (N) Tj 6 0 TD /F1 9 Tf -0.0395 Tc 0.7895 Tw (, where ) Tj 31.5 0 TD /F2 9 Tf -0.003 Tc 0 Tw (N) Tj 6 0 TD /F1 9 Tf -0.0176 Tc 1.3509 Tw ( is the total number of cells in the ) Tj -100.5 -12 TD 0.0009 Tc 2.9991 Tw (offspring group, termed propagule size ) Tj 3.003 Tc 0 Tw (\() Tj 160.5 0 TD /F2 9 Tf -0.003 Tc (N) Tj 6 0 TD /F1 9 Tf 0.0851 Tc 2.3149 Tw ( = 1 for zygote ) Tj -166.5 -12 TD 0.0411 Tc 1.3089 Tw (reproduction\). Evolution is defined as the change in adaptive features of a population over time as a result of natural selection Natural selection results in a process of adaptation , which means that, over generations, those features that are better adapted to the environment become more common This paper aims to introduce the reader to these new developments. 1997. In the transition) Tj 191.25 0 TD -0.0247 Tc 0.3372 Tw ( from single ) Tj -191.25 -11.25 TD 0.0256 Tc 1.2869 Tw (cells to multicellular organisms, the cells form cell groups ) Tj 0 -11.25 TD 0.1722 Tc 1.8903 Tw (belonging to the same species) Tj 129.75 4.5 TD /F1 6 Tf 0 Tc 0 Tw (6) Tj 3 -4.5 TD /F1 9.75 Tf 0.0066 Tc 1.3059 Tw (. This ) Tj T* 0.0989 Tc 3.6511 Tw (transfer of fitness from lower to higher levels occurs) Tj 0 Tc 0.5625 Tw ( ) Tj T* 0.2004 Tc 0.0621 Tw (through the evolution of cooperation an) Tj 162.75 0 TD -0.0018 Tc 0.0643 Tw (d conflict modifiers ) Tj -162.75 -11.25 TD 0.1766 Tc 1.1359 Tw (that restrict the opportunity for within group change and ) Tj 0 -11.25 TD 0.1759 Tc 0.7616 Tw (enhance the opportunity for between group change. Using a ) Tj T* -0.0684 Tc 0 Tw (multi) Tj 19.5 0 TD -0.2467 Tc (-) Tj 3 0 TD -0.0169 Tc 7.3294 Tw (level selection framework) Tj 115.5 4.5 TD /F1 6 Tf 0 Tc 0 Tw (1) Tj 3 0 TD 0.252 Tc (-) Tj 2.25 0 TD 0 Tc (5) Tj 3 -4.5 TD /F1 9.75 Tf 0.1561 Tc 6.5939 Tw (, it is possible to) Tj 0 Tc 0.5625 Tw ( ) Tj -146.25 -11.25 TD 0.1072 Tc 0.3719 Tw (understand how Darwin\222s properties emerge at a new level ) Tj 0 -11.25 TD -0.1385 Tc 0.951 Tw (\(for example, the mult) Tj 87.75 0 TD 0.0091 Tc 0.5534 Tw (icellular organism\) from the evolution ) Tj -87.75 -11.25 TD 0.0771 Tc 2.2354 Tw (of interactions in groups of lower level individuals \(for ) Tj 0 -11.25 TD 0.0411 Tc -0.1604 Tw (example, cells\). I have read many books about evolutionary biology in association with philosophical issues. Tj T* 0.1294 Tc -0.3169 Tw (For the group to become a new i) Tj 128.25 0 TD 0.0968 Tc -0.2843 Tw (ndividual, the organism, the ) Tj -128.25 -11.25 TD 0.213 Tc 0.8182 Tw (opportunity for between group change must be enhanced ) Tj 0 -11.25 TD 0.1411 Tc 2.0047 Tw (while the opportunity for within group change must be ) Tj T* 0.1475 Tc -0.335 Tw (restricted. 1999. ) The propagules then ) Tj T* -0.0273 Tc 1.2773 Tw (form offspring groups of the next generation \(dashed arrows\). ) We are interested in) Tj 0 Tc 0.5625 Tw ( ) Tj T* 0.1598 Tc 2.6527 Tw (understanding these first true emergent level functions ) Tj T* 0.032 Tc 5.7805 Tw (during the origin of the multicellular organism. What began as the study of animal social ) Tj 0 -11.25 TD 0.1513 Tc 0.0362 Tw (behavior, has now embraced the study of interactions at all ) Tj T* 0.0936 Tc 1.2189 Tw (biological levels. multi-level selection framework1-5, it is possible to understand how Darwin’s properties emerge at a new level (for example, the multicellular organism) from the evolution of interactions in groups of lower level individuals (for example, cells). An organism must have functions that maintain its ) Tj 0 -11.25 TD 0.1027 Tc 8.2098 Tw (integrity and individuality. Opposite Selection at two levels selection for at the level of the gene; against at the level of the genotype (organism). We believe that ) Tj T* 0.1059 Tc 0.5504 Tw (both kinds of mutations, selfish and uniformly deleterious, ) Tj T* 0.1415 Tc 7.421 Tw (are important for understanding the evolution of) Tj 0 Tc -0.1875 Tw ( ) Tj T* 0.2285 Tc 3.334 Tw (cooperation and fitness her) Tj 120.75 0 TD 0.0802 Tc 2.2323 Tw (itability during evolutionary ) Tj -120.75 -11.25 TD 0.1936 Tc 0.0877 Tw (transitions. History. Tj 107.25 0 TD 0 Tc -0.1875 Tw ( ) Tj -198 -11.25 TD -0.2813 Tc 0.0938 Tw (5. ) Those phenotypes differ because of genetic variation. It is as simple as Òsurvival of the Þttest.Ó There is, though, an important subtlety that is often omitted by the biology textbooks, providing a radical shift in the way we view evolution and natural selection. Tj 6.75 0 TD /F1 9 Tf -0.0649 Tc 1.0399 Tw ( in the models\). The author argues that the controversial concept of multilevel selection is essential for biological evolution, a suggestion that is made to stimulate new debate. Michaela Lewis . In the Evolution - Evolution - Types of selection: Natural selection can be studied by analyzing its effects on changing gene frequencies, but it can also be explored by examining its effects on the observable characteristics—or phenotypes—of individuals in a population. A multi-level approach of this kind allows important insights into the evolution of life, as well as bridging the long-standing conceptual chasm between molecular and organismal biologists. Tj -39.75 -11.25 TD 0.1232 Tc 3.3143 Tw (Under these conditions, an evolutionary transition will) Tj 0 Tc -0.1875 Tw ( ) Tj 0 -11.25 TD 0.1096 Tc 0 Tw (occur.) Each of these mechanisms of sexual selection has probably played a role over human evolution, but contest competition — the use Tj 78 0 TD /F2 9.75 Tf 0.1884 Tc 0.6241 Tw (Darwinian Dynamics, Evolutionary ) Tj -78 -11.25 TD 0.2322 Tc 3.3303 Tw (Transitions in Fitness and Individuality) Tj 177 0 TD /F1 9.75 Tf 0.0457 Tc 3.5168 Tw (. Tj 186 0 TD 0 Tc -0.1875 Tw ( ) Tj -162.75 -32.25 TD /F0 12 Tf -0.1836 Tc 0 Tw (Multi) Tj 27.75 0 TD -0.246 Tc (-) Tj 3.75 0 TD -0.1177 Tc -0.0698 Tw (level Selection, Cooperation and ) Tj 15.75 -15 TD -0.171 Tc 0.171 Tw (Variation of Fitness) Tj 98.25 0 TD 0 Tc 0 Tw ( ) Tj -168.75 -18 TD /F1 9.75 Tf 0.0669 Tc -0.0044 Tw (We take a multi) Tj 62.25 0 TD -0.2467 Tc 0 Tw (-) Tj 3 0 TD 0.1424 Tc -0.2049 Tw (level selection approach to fitness variation ) Tj -65.25 -11.25 TD 0.2016 Tc 0.3609 Tw (and evolutionary tr) Tj 78 0 TD 0.1131 Tc 0.1994 Tw (ansitions. Okasha’s book makes the sort of contribution that simply will not be able to be ignored by anyone interested in this field for many years to come. Summary: levels of selection • levels of selection • building higher levels • a levels of selection view of the world • genomic conflict • group selection 6 Levels of selection • I am pretending to a unity, deep inside myself, I know does not exist. Delet) Tj 105 0 TD 0.1017 Tc -0.1392 Tw (erious mutation may occur during ) Tj -105 -11.25 TD 0.1578 Tc -0.0953 Tw (cell division. BT 54 729 TD 0 0 0 rg /F1 9.75 Tf 0.0863 Tc 3.9048 Tw (cooperation of the lower level units. Evolution of Human Mate Choice David C. Geary, Jacob Vigil, and Jennifer Byrd-Craven University of Missouri – Columbia This article provides a review of evolutionary theory and empirical research on mate choices in non-human species and used as a frame for understanding the how and why of … Natural selection is one of the basic mechanisms of evolution, along with mutation, migration, and genetic drift. The author argues that the controversial concept of multilevel selection is essential for biological evolution, a suggestion that is made to stimulate new debate. #143 Summary of Selection and Evolution 1 Genetic variation within a population is the raw material on which natural selection can act. The basic theory was developed by both Darwin and Wallace, however, Darwin gave a much fuller argument. #143 Summary of Selection and Evolution 1 Genetic variation within a population is the raw material on which natural selection can act. Tj 102 0 TD 0 Tc -0.1875 Tw ( ) Tj -102 -11.25 TD ( ) Tj 9.75 0 TD 0.1261 Tc 1.3739 Tw (To be able to adapt to their environment, evolutionary) Tj 229.5 0 TD 0 Tc -0.1875 Tw ( ) Tj -239.25 -11.25 TD 0.1297 Tc 0.0995 Tw (individuals must be units of selection, and satisfy Darwin\222s ) Tj 0 -11.25 TD 0.1128 Tc 1.033 Tw (conditions of heritability and variation in fitness. It is crucial to distinguish the question concerning the level at which adaptations evolve from the question about the identity of the ultimate beneficiaries of that selection process. new work on the levels of selection, some of which has significantly re-defined the terms of the traditional debate. Tj 7.5 0 TD 0 Tc -0.1875 Tw ( ) Tj -111 -11.25 TD ( ) Tj 9.75 0 TD 0.0641 Tc 0.4984 Tw (In ) Tj 11.25 0 TD -0.1179 Tc 0 Tw (Figure) Tj 25.5 0 TD /F1 9 Tf 0 Tc ( ) Tj 3 0 TD (2) Tj 4.5 0 TD /F1 9.75 Tf 0.097 Tc -0.2845 Tw (, the two different species \(h) Tj 113.25 0 TD 0.2204 Tc -0.4079 Tw (ost and symbiont, ) Tj -167.25 -11.25 TD 0.1247 Tc 1.094 Tw (predator and prey, two different syntrophic feeders, etc.\), ) Tj 0 -11.25 TD 0.1692 Tc 0.9766 Tw (are indicated by squares and circles. Professor Samir Okasha's book "Evolution and the Levels of Selection" is a materpiece that is second to none! There will also be a change in gene frequency in ) Tj -42.75 -11.25 TD 0.1617 Tc 0.2341 Tw (the global population due to differences in fitness between ) Tj 0 -11.25 TD 0.1997 Tc -0.106 Tw (the adult groups \(dotted arrows\). The question of levels of selection - on which biologists and philosophers have long disagreed - is central to evolutionary theory and to the philosophy of biology. 1999.) Tj 9 0 TD 0 Tc -0.1875 Tw ( ) Tj 4.5 0 TD -0.0989 Tc 3.0614 Tw (Michod, R.E. Tj 189 0 TD 0 Tc -0.1875 Tw ( ) Tj -189 -11.25 TD ( ) Tj 9.75 0 TD 0.1095 Tc 2.1405 Tw (Cooperation is fundamental to the emergence of new ) Tj -9.75 -11.25 TD 0.0975 Tc 4.515 Tw (levels of fitness in the biological) Tj 0 Tc -0.1875 Tw ( ) Tj 159.75 0 TD 0.1552 Tc 4.1573 Tw (hierarchy, because) Tj 0 Tc -0.1875 Tw ( ) Tj -159.75 -11.25 TD 0.1983 Tc 1.5308 Tw (cooperation increases the fitness of the group, and new ) Tj 0 -11.25 TD 0.1863 Tc 2.0262 Tw (units of selection start out as groups of existing units. ) We have studied deleterious mutations using a ) Tj 0 -11.25 TD 0.169 Tc 2.965 Tw (variety of deterministic and stochastic techniques that ) Tj T* 0.1008 Tc 0.3867 Tw (allow us to study a distribution of mutational effects \(from ) Tj 0.0706 0.0784 0.0824 rg ET 69.75 729.75 m 69 729.75 l 69 729 l 69.75 729.75 l 69 729 l 69.75 729 l 69.75 729.75 l h f 69 729.75 m 69.75 729.75 l 69 729.75 l h f 69.75 729.75 m 69 729.75 l 69.75 729.75 l h f 69.75 729.75 m h f 69.75 729.75 m h f 70.5 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77.25 727.5 m h f 77.25 727.5 m h f 77.25 726.75 m 77.25 727.5 l 77.25 726.75 l h f 77.25 726.75 m 77.25 727.5 l 77.25 726.75 l h f 77.25 726.75 m h f 77.25 726.75 m h f 77.25 727.5 m 77.25 726.75 l 77.25 727.5 l h f 77.25 728.25 m 77.25 727.5 l 77.25 728.25 l h f 76.5 729 m 77.25 728.25 l 76.5 729 l h f 76.5 729 m h f 76.5 729.75 m 75.75 729.75 l 76.5 729 l 76.5 729.75 l h f 75.75 730.5 m 75.75 729.75 l 76.5 729.75 l 75.75 730.5 l h f 75.75 730.5 m 75.75 729.75 l 75.75 730.5 l h f 75.75 730.5 m 75 730.5 l 75.75 730.5 l h f 75 731.25 m 75 730.5 l 75.75 730.5 l 75 731.25 l h f 75 731.25 m h f 75 731.25 m h f 75 732 m 75 731.25 l 75 732 l h f 74.25 732 m 75 732 l 74.25 732 l h f 74.25 732 m h f 74.25 732.75 m 74.25 732 l 74.25 732.75 l 74.25 732 l 74.25 732.75 l h f 74.25 732 m 74.25 732.75 l 74.25 732 l h f 73.5 732.75 m 74.25 732 l 74.25 732.75 l 73.5 732.75 l h f 73.5 732.75 m 74.25 732.75 l 73.5 732.75 l 74.25 732.75 l 73.5 732.75 l h f 73.5 732.75 m h f 73.5 732.75 m h f 73.5 733.5 m 73.5 732.75 l 73.5 733.5 l 73.5 732.75 l 73.5 733.5 l h f 72.75 732.75 m 73.5 732.75 l 73.5 733.5 l 72.75 732.75 l h f 73.5 733.5 m 73.5 732.75 l 73.5 733.5 l h f 72.75 733.5 m 72.75 732.75 l 73.5 733.5 l 72.75 733.5 l h f 72.75 733.5 m h f 72.75 733.5 m h f 72.75 733.5 m h f 72.75 733.5 m h f 72.75 733.5 m h f 72 733.5 m 72.75 733.5 l 72 733.5 l h f 72 733.5 m 72.75 733.5 l 72 733.5 l 72.75 733.5 l 72 733.5 l h f 72 733.5 m h f 72 733.5 m h f 72.75 733.5 m 72 733.5 l 72.75 733.5 l h f 72.75 733.5 m h f 73.5 733.5 m 72.75 733.5 l 73.5 733.5 l h f 73.5 733.5 m h f 73.5 733.5 m h f 74.25 733.5 m 73.5 733.5 l 74.25 733.5 l h f 74.25 733.5 m 73.5 733.5 l 74.25 733.5 l h f 74.25 733.5 m h f 74.25 733.5 m h f 74.25 733.5 m 74.25 732.75 l 74.25 733.5 l h f 75 732.75 m 74.25 733.5 l 74.25 732.75 l 75 732.75 l h f 75 732.75 m h f 75 732.75 m h f 75 732 m 75 732.75 l 75 732 l h f 75 732 m 75.75 732 l 75 732.75 l 75 732 l h f 75.75 732 m 75 732 l 75.75 732 l h f 75.75 732 m h f 75.75 732 m h f 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665.25 m h f 112.5 665.25 m 113.25 665.25 l 112.5 665.25 l h f 112.5 665.25 m 113.25 665.25 l 112.5 665.25 l h f 112.5 664.5 m 112.5 665.25 l 112.5 664.5 l h f 112.5 664.5 m 112.5 665.25 l 112.5 664.5 l 112.5 665.25 l 112.5 664.5 l h f 112.5 664.5 m h f 112.5 664.5 m 111.75 664.5 l 112.5 664.5 l h f 111.75 664.5 m 111.75 663.75 l 112.5 664.5 l 111.75 664.5 l h f 111.75 664.5 m h f 111.75 663.75 m 111.75 664.5 l 111.75 663.75 l h f 111.75 663.75 m h f 111 663.75 m 111.75 663.75 l 111 663.75 l h f 111.75 663.75 m 111 663.75 l 111.75 663.75 l h f 111 664.5 m 111 663.75 l 111.75 663.75 l 111 664.5 l h f 111 664.5 m h f 111 664.5 m h f 111 665.25 m 111 664.5 l 111 665.25 l h f 111 665.25 m h f 111 666 m 111 665.25 l 111 666 l h f 111.75 666 m 111 666 l 111.75 666 l h f 111 666 m 111.75 666 l 111 666 l h f 111.75 666 m 111.75 666.75 l 111 666 l 111.75 666 l h f 111.75 666.75 m 111.75 666 l 111.75 666.75 l h f 112.5 666.75 m 111.75 666.75 l 112.5 666.75 l h f 112.5 667.5 m 111.75 666.75 l 112.5 666.75 l 112.5 667.5 l h f 112.5 667.5 m 112.5 666.75 l 112.5 667.5 l h f 112.5 667.5 m h f 113.25 667.5 m 112.5 667.5 l 113.25 667.5 l h f 113.25 667.5 m h f 113.25 668.25 m 113.25 667.5 l 113.25 668.25 l h f 114 667.5 m 114 668.25 l 113.25 668.25 l 113.25 667.5 l 114 667.5 l h f 114 668.25 m 113.25 668.25 l 114 667.5 l 114 668.25 l h f 114 668.25 m 114.75 668.25 l 114 668.25 l h f 114.75 668.25 m 114 668.25 l 114.75 668.25 l h f 114.75 668.25 m h f 115.5 668.25 m 114.75 668.25 l 115.5 668.25 l h f 115.5 669 m 114.75 668.25 l 115.5 668.25 l 115.5 669 l h f 116.25 668.25 m 116.25 669 l 115.5 669 l 115.5 668.25 l 116.25 668.25 l h f 116.25 668.25 m h f 116.25 668.25 m 116.25 669 l 116.25 668.25 l h f 116.25 668.25 m 116.25 669 l 116.25 668.25 l h f 117 669 m 116.25 669 l 116.25 668.25 l 117 668.25 l 117 669 l h f 117 668.25 m h f 117 668.25 m 117 669 l 117 668.25 l h f 117 668.25 m h f 117 668.25 m h f 117 669 m 117 668.25 l 117 669 l h f 117 669 m h f 117 669 m h f 117 669 m h f 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108.75 668.25 l h f 108 668.25 m 108.75 668.25 l 108 668.25 l h f 108 668.25 m h f 108 667.5 m 108 668.25 l 108 667.5 l h f 107.25 668.25 m 107.25 667.5 l 108 667.5 l 108 668.25 l 107.25 668.25 l h f 107.25 667.5 m 108 667.5 l 107.25 668.25 l 107.25 667.5 l h f 107.25 667.5 m h f 106.5 667.5 m 107.25 667.5 l 106.5 667.5 l h f 106.5 667.5 m 107.25 667.5 l 106.5 667.5 l h f 106.5 667.5 m 107.25 667.5 l 106.5 667.5 l h f 106.5 666.75 m 106.5 667.5 l 106.5 666.75 l h f 106.5 667.5 m 106.5 666.75 l 106.5 667.5 l h f 106.5 667.5 m h f 106.5 667.5 m h f 106.5 668.25 m 106.5 667.5 l 106.5 668.25 l h f 106.5 668.25 m 106.5 667.5 l 106.5 668.25 l h f 106.5 668.25 m h f 107.25 668.25 m 106.5 669 l 106.5 668.25 l 107.25 668.25 l h f 107.25 669 m 106.5 669 l 107.25 668.25 l 107.25 669 l h f 107.25 669 m h f 107.25 669 m 108 669 l 107.25 669 l h f 108 669 m 107.25 669 l 108 669 l h f 108 669.75 m 108 669 l 108 669.75 l h f 108.75 669.75 m 108 669.75 l 108.75 669.75 l h f 108.75 669.75 m h f 109.5 670.5 m 108.75 670.5 l 108.75 669.75 l 109.5 669.75 l 109.5 670.5 l h f 109.5 669.75 m 109.5 670.5 l 109.5 669.75 l h f 109.5 670.5 m 109.5 669.75 l 109.5 670.5 l h f 110.25 670.5 m 109.5 670.5 l 110.25 670.5 l h f 110.25 670.5 m h f 111 670.5 m 110.25 670.5 l 111 670.5 l h f 111 671.25 m 111 670.5 l 111.75 670.5 l 111 671.25 l h f 111.75 670.5 m 111.75 671.25 l 111 671.25 l 111 670.5 l 111.75 670.5 l h f 111.75 671.25 m 111.75 670.5 l 111.75 671.25 l h f 112.5 671.25 m 111.75 671.25 l 112.5 671.25 l h f 112.5 671.25 m h f 113.25 671.25 m 112.5 671.25 l 113.25 671.25 l h f 113.25 671.25 m h f 114 671.25 m 113.25 671.25 l 114 671.25 l h f 114 671.25 m h f 114 671.25 m h f 114 671.25 m h f 114.75 671.25 m 114 671.25 l 114.75 671.25 l h f 113.25 671.25 m 114.75 671.25 l 113.25 671.25 l h f 113.25 672 m 113.25 671.25 l 114 671.25 l 113.25 672 l h f 112.5 672 m 113.25 672 l 112.5 672 l h f 111.75 672.75 m 112.5 672 l 111.75 672.75 l h f 111 672.75 m 111.75 672 l 111.75 672.75 l 111 672.75 l h f 110.25 672.75 m 111 672.75 l 110.25 672.75 l h f 110.25 673.5 m 110.25 672.75 l 110.25 673.5 l h f 109.5 672.75 m 110.25 672.75 l 110.25 673.5 l 109.5 673.5 l 109.5 672.75 l h f 109.5 673.5 m 108.75 673.5 l 109.5 672.75 l 109.5 673.5 l h f 108.75 674.25 m 108.75 673.5 l 109.5 673.5 l 108.75 674.25 l h f 108 673.5 m 108.75 673.5 l 108.75 674.25 l 108 674.25 l 108 673.5 l h f 108 674.25 m 108 673.5 l 108 674.25 l h f 107.25 674.25 m 108 674.25 l 107.25 674.25 l h f 107.25 674.25 m h f 106.5 674.25 m 107.25 674.25 l 106.5 674.25 l h f 106.5 674.25 m 105.75 674.25 l 106.5 674.25 l h f 105.75 674.25 m 106.5 674.25 l 105.75 674.25 l h f 105.75 674.25 m h f 105 675 m 105 674.25 l 105.75 674.25 l 105 675 l h f 105 675 m 105 674.25 l 105 675 l h f 104.25 675 m 104.25 674.25 l 105 674.25 l 105 675 l 104.25 675 l h f 104.25 674.25 m 104.25 675 l 104.25 674.25 l h f 104.25 675 m 104.25 674.25 l 104.25 675 l h f 104.25 675 m h f 103.5 675 m 104.25 675 l 103.5 675 l h f 103.5 675 m h f 103.5 675 m h f 102.75 675 m 103.5 675 l 102.75 675 l h f 102.75 675 m h f 102.75 675 m h f 102.75 675 m h f 102 675 0.75 0.24 re f BT 1.0111 0 0 1 133.5 591.75 Tm /F3 8.961 Tf -0.0647 Tc 0.5398 Tw (offspring group) Tj 11.8682 -12 TD -0.0746 Tc 10.9343 Tw (\(size ) Tj 11.1094 Tc 0 Tw (\)) Tj ET BT 1.0112 0 0 1 167.25 579.75 Tm /F4 11.0904 Tf 0.1515 Tc (N) Tj ET BT 1.0108 0 0 1 175.5 579.75 Tm /F4 8.961 Tf 0 Tc -0.0143 Tw ( ) Tj ET BT 1.0111 0 0 1 94.5 597 Tm /F3 8.961 Tf -0.2659 Tw ( ) Tj ET 63 678.75 m 62.25 678 l 60 678 l 59.25 678.75 l 59.25 678.75 l 59.25 679.5 l 60 679.5 l 60 680.25 l 61.5 680.25 l 61.5 681 l 62.25 681 l 62.25 681.75 l 60 681.75 l 60 681 l 59.25 681 l 59.25 681.75 l 60 681.75 l 60 682.5 l 63 682.5 l 63 678.75 l 63.75 678.75 l 63.75 678 l 63 678 l 63 678.75 l 62.25 680.25 m 61.5 680.25 l 61.5 679.5 l 60 679.5 l 60 678.75 l 60.75 678.75 l 60.75 678 l 61.5 678 l 61.5 678.75 l 62.25 678.75 l 62.25 680.25 l h f* 67.5 678 m 67.5 678.75 l 67.5 678 l 65.25 678 l 65.25 678.75 l 64.5 678.75 l 64.5 679.5 l 64.5 679.5 l 64.5 681.75 l 65.25 681.75 l 65.25 682.5 l 67.5 682.5 l 67.5 681.75 l 67.5 684 l 68.25 684 l 68.25 678 l 67.5 678 l 65.25 680.25 m 65.25 678.75 l 66 678.75 l 66.75 678 l 66.75 678.75 l 67.5 678.75 l 67.5 681.75 l 65.25 681.75 l 65.25 680.25 l h f* 72.75 678 m 72.75 678.75 l 72.75 678 l 69.75 678 l 69.75 678.75 l 69.75 678.75 l 69.75 682.5 l 70.5 682.5 l 70.5 678.75 l 72.75 678.75 l 72.75 682.5 l 73.5 682.5 l 73.5 678 l 72.75 678 l h f* 74.25 678 m 74.25 684 l 75 684 l 75 678 l 74.25 678 l h f* 78 678.75 m 78.75 678 l 77.25 678 l 76.5 678.75 l 76.5 678.75 l 76.5 684 l 77.25 684 l 77.25 682.5 l 78 682.5 l 78 681.75 l 77.25 681.75 l 77.25 678.75 l 78 678.75 l h f* 81.75 677.25 m 82.5 677.25 l 82.5 676.5 l 84 676.5 l 84 677.25 l 84 677.25 l 84.75 677.25 l 84.75 678.75 l 84.75 678 l 81.75 678 l 81.75 679.5 l 81 679.5 l 81 681 l 81.75 681 l 81.75 682.5 l 84.75 682.5 l 84.75 681.75 l 84.75 682.5 l 85.5 682.5 l 85.5 676.5 l 84.75 676.5 l 84 675.75 l 82.5 675.75 l 82.5 676.5 l 81.75 676.5 l 81.75 677.25 l 82.5 680.25 m 82.5 678.75 l 84.75 678.75 l 84.75 681 l 84 681.75 l 82.5 681.75 l 82.5 680.25 l h f* 86.25 678 m 86.25 682.5 l 87 682.5 l 87 681.75 l 87.75 681.75 l 87.75 682.5 l 89.25 682.5 l 89.25 681.75 l 87.75 681.75 l 87.75 681 l 87 681 l 87 678 l 86.25 678 l h f* 89.25 680.25 m 89.25 681.75 l 90 681.75 l 90 682.5 l 93 682.5 l 93 681.75 l 93.75 681.75 l 93.75 681 l 93.75 678.75 l 93 678.75 l 93 678 l 90 678 l 90 678.75 l 89.25 678.75 l 89.25 680.25 l 90 680.25 m 90 678.75 l 91.5 678.75 l 91.5 678 l 91.5 678.75 l 92.25 678.75 l 93 679.5 l 93 681 l 92.25 681 l 92.25 681.75 l 90.75 681.75 l 90 681 l 90 680.25 l h f* 97.5 678 m 97.5 678.75 l 97.5 678 l 94.5 678 l 94.5 678.75 l 94.5 678.75 l 94.5 682.5 l 95.25 682.5 l 95.25 678.75 l 97.5 678.75 l 97.5 682.5 l 98.25 682.5 l 98.25 678 l 97.5 678 l h f* 99.75 675.75 m 99.75 682.5 l 100.5 682.5 l 100.5 681.75 l 100.5 682.5 l 102.75 682.5 l 102.75 682.5 l 102.75 681.75 l 103.5 681.75 l 103.5 678.75 l 102.75 678.75 l 102.75 678 l 100.5 678 l 100.5 675.75 l 99.75 675.75 l 100.5 680.25 m 100.5 678.75 l 101.25 678.75 l 101.25 678 l 102 678.75 l 102.75 678.75 l 102.75 681.75 l 100.5 681.75 l 100.5 680.25 l h f* 69 667.5 m 69.75 667.5 l 69.75 666.75 l 72 666.75 l 72 666.75 l 72 667.5 l 71.25 667.5 l 71.25 668.25 l 69.75 668.25 l 69 669 l 69 670.5 l 72 670.5 l 72 669.75 l 72.75 669.75 l 72 669 l 72 669.75 l 71.25 669.75 l 71.25 670.5 l 70.5 670.5 l 70.5 669.75 l 69.75 669.75 l 69.75 669 l 71.25 669 l 72 668.25 l 72.75 668.25 l 72.75 666.75 l 72 666.75 l 72 666 l 69 666 l 69 667.5 l h f* 73.5 671.25 m 73.5 672.75 l 74.25 672.75 l 74.25 671.25 l 73.5 671.25 l 73.5 666 m 73.5 670.5 l 74.25 670.5 l 74.25 666 l 73.5 666 l h f* 75 666 m 75 666.75 l 78 669.75 l 75.75 669.75 l 75.75 670.5 l 79.5 670.5 l 76.5 667.5 l 76.5 666.75 l 79.5 666.75 l 79.5 666 l 75 666 l h f* 83.25 667.5 m 84 667.5 l 84 666.75 l 83.25 666.75 l 83.25 666 l 81 666 l 80.25 666.75 l 80.25 666.75 l 80.25 670.5 l 84 670.5 l 84 668.25 l 81 668.25 l 81 666.75 l 83.25 666.75 l 83.25 667.5 l 81 669 m 83.25 669 l 83.25 669.75 l 82.5 669.75 l 82.5 670.5 l 81.75 670.5 l 81.75 669.75 l 81 669.75 l 81 669 l h f* 90 673.5 m 88.5 668.25 l 89.25 669 l 90 669 l 90 669.75 l 90.75 669.75 l 90.75 670.5 l 90.75 670.5 l 90 670.5 l 92.25 670.5 l 90.75 670.5 l 90.75 669.75 l 90 669 l 90 667.5 l 90.75 667.5 l 90.75 666.75 l 91.5 666.75 l 91.5 667.5 l 91.5 666 l 90 666 l 90 667.5 l 89.25 667.5 l 89.25 668.25 l 88.5 668.25 l 87.75 666 l 87 666 l 89.25 672 l 89.25 673.5 l 88.5 673.5 l 90 673.5 l h f* 93.75 671.25 m 94.5 670.5 l 94.5 669.75 l 94.5 669.75 l 93.75 669.75 l 93.75 671.25 l 93.75 669 m 93 665.25 l 93 664.5 l 92.25 664.5 l 92.25 663.75 l 91.5 663.75 l 91.5 663 l 90.75 663 l 90.75 663.75 l 90.75 663 l 91.5 663 l 91.5 663.75 l 92.25 663.75 l 92.25 664.5 l 93 667.5 l 93 669 l 92.25 669 l 93.75 669 l h f* BT 1.0111 0 0 1 58.5 635.25 Tm -0.011 Tc 0 Tw (development) Tj 0.7418 -9.75 TD 0.0128 Tc (within-group) Tj 9.6429 -10.5 TD 0.0463 Tc (change) Tj 20.0275 54 TD 0 Tc -0.2659 Tw ( ) Tj 1 1 1 rg ET 128.25 593.25 m 127.5 592.5 l 127.5 591.75 l 126.75 591.75 l 126.75 591 l 126 591 l 125.25 590.25 l 124.5 590.25 l 123.75 589.5 l 120.75 589.5 l 120 588.75 l 117.75 588.75 l 117 589.5 l 114 589.5 l 113.25 590.25 l 112.5 590.25 l 111.75 591 l 111 591 l 111 591.75 l 110.25 591.75 l 110.25 592.5 l 109.5 593.25 l 109.5 594 l 110.25 594.75 l 110.25 595.5 l 111 595.5 l 111 596.25 l 111.75 596.25 l 112.5 597 l 113.25 597 l 114 597.75 l 116.25 597.75 l 117 598.5 l 120.75 598.5 l 121.5 597.75 l 123.75 597.75 l 124.5 597 l 125.25 597 l 126 596.25 l 126.75 596.25 l 126.75 595.5 l 127.5 595.5 l 127.5 594.75 l 128.25 594 l 128.25 593.25 l h f* 0.0706 0.0784 0.0824 rg 119.25 588.75 m 121.5 588.75 l 122.25 589.5 l 124.5 589.5 l 125.25 590.25 l 126 590.25 l 128.25 592.5 l 128.25 593.25 l 127.5 593.25 l 127.5 592.5 l 125.25 590.25 l 124.5 590.25 l 123.75 590.25 l 123 589.5 l 119.25 589.5 l h f 109.5 593.25 m 109.5 592.5 l 111.75 590.25 l 112.5 590.25 l 113.25 589.5 l 115.5 589.5 l 116.25 588.75 l 119.25 588.75 l 119.25 589.5 l 114.75 589.5 l 114 590.25 l 112.5 590.25 l 111.75 591 l 111.75 591 l 110.25 592.5 l 110.25 593.25 l h f 119.25 598.5 m 115.5 598.5 l 114.75 597.75 l 113.25 597.75 l 112.5 597 l 111.75 597 l 110.25 595.5 l 110.25 594.75 l 109.5 594.75 l 109.5 593.25 l 110.25 593.25 l 110.25 594.75 l 112.5 597 l 113.25 597 l 114 597 l 114.75 597.75 l 119.25 597.75 l h f 128.25 593.25 m 128.25 594.75 l 127.5 594.75 l 127.5 595.5 l 126 597 l 125.25 597 l 124.5 597.75 l 123 597.75 l 122.25 598.5 l 119.25 598.5 l 119.25 597.75 l 123 597.75 l 123.75 597 l 125.25 597 l 126 596.25 l 126 596.25 l 127.5 594.75 l 127.5 593.25 l h f 1 1 1 rg 85.5 699 m 83.25 696.75 l 82.5 696.75 l 81 696 l 75.75 696 l 75 696.75 l 73.5 696.75 l 72 698.25 l 72 700.5 l 73.5 702 l 75 702 l 75.75 702.75 l 81 702.75 l 82.5 702 l 83.25 702 l 85.5 699.75 l 85.5 699 l h f* 0.0706 0.0784 0.0824 rg 78.75 696 m 81 696 l 82.5 696.75 l 83.25 696.75 l 84.75 698.25 l 85.5 698.25 l 85.5 699 l 84.75 699 l 84.75 698.25 l 84 697.5 l 83.25 697.5 l 82.5 696.75 l 81 696.75 l 80.25 696 l 78.75 696 l h f 72 699 m 72 698.25 l 73.5 696.75 l 75 696.75 l 75.75 696 l 78.75 696 l 77.25 696 l 75.75 696.75 l 75 696.75 l 73.5 697.5 l 72.75 697.5 l 72.75 698.25 l 72 698.25 l 72 699 l h f 78.75 702.75 m 75.75 702.75 l 75 702 l 73.5 702 l 72 700.5 l 72 699 l 72 700.5 l 72.75 700.5 l 72.75 701.25 l 73.5 701.25 l 75 702 l 75.75 702 l 77.25 702.75 l 78.75 702.75 l h f 85.5 699 m 85.5 700.5 l 84.75 700.5 l 83.25 702 l 82.5 702 l 81 702.75 l 78.75 702.75 l 80.25 702.75 l 81 702 l 82.5 702 l 83.25 701.25 l 84 701.25 l 84.75 700.5 l 84.75 699 l h f 1 1 1 rg 230.25 706.5 m 229.5 705 l 229.5 702 l 228.75 700.5 l 228.75 699 l 227.25 697.5 l 226.5 696 l 225 694.5 l 224.25 693 l 222 690.75 l 220.5 690 l 219.75 689.25 l 218.25 688.5 l 217.5 687.75 l 214.5 686.25 l 213.75 685.5 l 212.25 685.5 l 209.25 684 l 207.75 684 l 206.25 683.25 l 204.75 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723.75 l 218.25 724.5 l 217.5 725.25 l 214.5 726.75 l 213.75 727.5 l 212.25 727.5 l 209.25 729 l 207.75 729 l 206.25 729 l 204.75 729.75 l 198.75 729.75 l 203.25 729.75 l 204.75 729 l 207.75 729 l 209.25 728.25 l 210.75 728.25 l 213.75 726.75 l 214.5 726.75 l 217.5 725.25 l 218.25 724.5 l 219.75 723.75 l 220.5 723 l 222 722.25 l 223.5 720.75 l 224.25 719.25 l 225.75 717.75 l 226.5 716.25 l 228 714.75 l 228.75 713.25 l 228.75 712.5 l 229.5 711 l 229.5 706.5 l h f 258.75 650.25 m 258.75 649.5 l 258.75 650.25 l h f 258.75 649.5 m h f 259.5 650.25 m 258.75 650.25 l 258.75 649.5 l 259.5 650.25 l h f 259.5 650.25 m h f 259.5 650.25 m h f 260.25 650.25 m 259.5 650.25 l 260.25 650.25 l h f 260.25 650.25 m h f 260.25 650.25 m h f 260.25 650.25 m h f 261 650.25 m 260.25 650.25 l 261 650.25 l h f 261 650.25 m h f 261.75 649.5 m 261.75 650.25 l 261 650.25 l 261.75 649.5 l h f 261.75 650.25 m 261.75 649.5 l 261.75 650.25 l h f 262.5 649.5 m 261.75 650.25 l 261.75 649.5 l 262.5 649.5 l h f 262.5 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281.25 693.75 m h f 281.25 693.75 m h f 280.5 693.75 m 281.25 693.75 l 280.5 693.75 l h f 280.5 693.75 m h f 280.5 693.75 m h f 280.5 693.75 m h f 280.5 693.75 m h f 280.5 693 m 280.5 693.75 l 280.5 693 l 280.5 693.75 l h f 280.5 693.75 m 280.5 693 l 280.5 693.75 l h f 279.75 693.75 m 280.5 693 l 280.5 693.75 l 279.75 693.75 l 280.5 693.75 l 279.75 693.75 l h f 280.5 693 m 280.5 693.75 l 279.75 693.75 l 279.75 693 l 280.5 693 l h f 279.75 693 m 280.5 693 l 279.75 693 l h f 279.75 693 m h f 279.75 693 m h f 279.75 693 m h f 279.75 692.25 m 279.75 693 l 279.75 692.25 l h f 279.75 693 m h f 279.75 693 m h f 279 692.25 m 279.75 692.25 l 279.75 693 l 279 692.25 l h f 279 692.25 m h f 279 692.25 m 279.75 692.25 l 279 692.25 l 279.75 692.25 l h f 279 692.25 m h f 279 692.25 m h f 279 692.25 m h f 279 692.25 m h f 279 691.5 m 279 692.25 l 279 691.5 l h f 278.25 691.5 m 279 691.5 l 279 692.25 l 278.25 691.5 l 279 692.25 l 278.25 692.25 l 278.25 691.5 l h f 278.25 691.5 m 279 691.5 l 278.25 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f 280.5 693.75 m 280.5 694.5 l 280.5 693.75 l h f 280.5 693.75 m h f 281.25 693.75 m 281.25 694.5 l 280.5 694.5 l 280.5 693.75 l 281.25 693.75 l h f 281.25 693.75 m 281.25 694.5 l 281.25 693.75 l h f 281.25 693.75 m h f 281.25 693.75 m h f 282 693.75 m 281.25 694.5 l 281.25 693.75 l 282 693.75 l 281.25 693.75 l 282 693.75 l h f 282 693.75 m 281.25 693.75 l 282 693.75 l 281.25 693.75 l 282 693.75 l h f 282 694.5 m 281.25 694.5 l 281.25 693.75 l 282 693.75 l 282 694.5 l h f 282 694.5 m 281.25 694.5 l 282 694.5 l h f 281.25 694.5 m 282 694.5 l 281.25 694.5 l 282 694.5 l 281.25 694.5 l h f 281.25 694.5 m h f 281.25 694.5 m h f 280.5 694.5 m 281.25 694.5 l 280.5 694.5 l h f 280.5 694.5 m h f 280.5 694.5 m h f 280.5 694.5 m h f 279.75 694.5 m 280.5 694.5 l 279.75 694.5 l h f 279.75 694.5 m h f 279.75 694.5 m h f 279 694.5 m 279.75 694.5 l 279 694.5 l h f 279 694.5 m h f 279 694.5 m h f 278.25 694.5 m 279 694.5 l 278.25 694.5 l h f 278.25 694.5 m 279 694.5 l 278.25 694.5 l h f 278.25 694.5 m 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693.75 l h f 276 694.5 m 276 693.75 l 276 694.5 l h f 276 694.5 m h f 276.75 694.5 m 276 694.5 l 276.75 694.5 l h f 276.75 694.5 m h f 276.75 694.5 m 276 694.5 l 276.75 694.5 l h f 276.75 694.5 m h f 276.75 695.25 m 276.75 694.5 l 276.75 695.25 l h f 276.75 694.5 m 276.75 695.25 l 276.75 694.5 l h f 277.5 695.25 m 276.75 695.25 l 276.75 694.5 l 277.5 694.5 l 277.5 695.25 l h f 277.5 695.25 m h f 277.5 695.25 m h f 278.25 695.25 m 277.5 695.25 l 278.25 695.25 l h f 278.25 695.25 m h f 278.25 695.25 m h f 279 695.25 m 278.25 695.25 l 279 695.25 l h f 279 695.25 m h f 279 695.25 m h f 279 695.25 m h f 279 696 m 279 695.25 l 279 696 l h f 279.75 695.25 m 279.75 696 l 279 696 l 279 695.25 l 279.75 695.25 l h f 279.75 695.25 m 279.75 696 l 279.75 695.25 l h f 279.75 695.25 m 279.75 696 l 279.75 695.25 l h f 279.75 696 m 279.75 695.25 l 279.75 696 l h f 279.75 695.25 m 279.75 696 l 279.75 695.25 l h f 279.75 695.25 m h f 279.75 696 m 279.75 695.25 l 279.75 696 l h f 279 696 m 279.75 695.25 l 279.75 696 l 279 696 l h f 279 696 m h f 279 696.75 m 278.25 696 l 279 696 l 279 696.75 l h f 278.25 696 m 279 696 l 279 696.75 l 278.25 696.75 l 278.25 696 l h f 277.5 696.75 m 278.25 696 l 278.25 696.75 l 277.5 696.75 l h f 277.5 696.75 m 278.25 696.75 l 277.5 696.75 l h f 277.5 696.75 m h f 276.75 696.75 m 277.5 696.75 l 276.75 696.75 l h f 276.75 696.75 m h f 276.75 697.5 m 276.75 696.75 l 276.75 697.5 l h f 276.75 697.5 m 276.75 696.75 l 276.75 697.5 l h f 276 697.5 m 276 696.75 l 276.75 696.75 l 276.75 697.5 l 276 697.5 l h f 276 697.5 m 276 696.75 l 276 697.5 l h f 276 697.5 m 275.25 697.5 l 276 697.5 l h f 275.25 697.5 m 276 697.5 l 275.25 697.5 l h f 275.25 697.5 m h f 275.25 697.5 m h f 275.25 697.5 m h f 274.5 697.5 m 275.25 697.5 l 274.5 697.5 l h f 274.5 697.5 m h f 274.5 697.5 m h f 274.5 697.5 m h f 273.75 697.5 m 274.5 697.5 l 273.75 697.5 l h f 273.75 697.5 m h f 273.75 697.5 m h f 273.75 697.5 m h f 273.75 697.5 m h f 273.75 697.5 m h f 273.75 697.5 m h f 273.75 697.5 m h f 273 697.5 0.75 0.24 re f 282.75 681.75 m 282.75 667.5 l 283.5 667.5 l 283.5 681.75 l 282.75 681.75 l h f* 282.75 687 m 280.5 681.75 l 281.25 681.75 l 281.25 682.5 l 284.25 682.5 l 284.25 681.75 l 285 681.75 l 282.75 687 l h f* 1 1 1 rg 289.5 662.25 m 289.5 660.75 l 288.75 660.75 l 288.75 660 l 287.25 659.25 l 286.5 659.25 l 285.75 658.5 l 280.5 658.5 l 279 659.25 l 278.25 659.25 l 276.75 660 l 276.75 660.75 l 276 660.75 l 276 663 l 276.75 663 l 276.75 663.75 l 277.5 664.5 l 278.25 664.5 l 279 664.5 l 280.5 665.25 l 285.75 665.25 l 286.5 664.5 l 288 664.5 l 288.75 663.75 l 288.75 663 l 289.5 663 l 289.5 662.25 l h f* 0.0706 0.0784 0.0824 rg 282.75 658.5 m 285.75 658.5 l 286.5 659.25 l 288 659.25 l 289.5 660.75 l 289.5 662.25 l 289.5 661.5 l 288 660 l 287.25 660 l 286.5 659.25 l 285.75 659.25 l 284.25 658.5 l 282.75 658.5 l h f 276 662.25 m 276 660.75 l 276.75 660 l 278.25 659.25 l 279 659.25 l 280.5 658.5 l 282.75 658.5 l 281.25 658.5 l 280.5 659.25 l 279 659.25 l 278.25 660 l 277.5 660 l 276 661.5 l 276 662.25 l h f 282.75 666 m 281.25 666 l 280.5 665.25 l 279 665.25 l 278.25 664.5 l 277.5 664.5 l 276.75 663.75 l 276 663.75 l 276 662.25 l 276.75 663 l 276.75 663.75 l 277.5 663.75 l 278.25 664.5 l 279 664.5 l 280.5 665.25 l 282.75 665.25 l h f 289.5 662.25 m 289.5 663.75 l 288.75 663.75 l 288 664.5 l 286.5 665.25 l 285.75 665.25 l 284.25 666 l 282.75 666 l 282.75 665.25 l 285.75 665.25 l 286.5 664.5 l 287.25 664.5 l 288 663.75 l 288.75 663.75 l 288.75 663 l 289.5 662.25 l h f q 179.25 706.5 111 11.25 re h W n BT 1.0111 0 0 1 179.25 708.75 Tm -0.142 Tc 0 Tw (propagule) Tj ET Q BT 1.0111 0 0 1 219 708.75 Tm ( ) Tj -28.1869 -10.5 TD -0.1544 Tc 0 Tw (pool) Tj 17.0605 0 TD 0 Tc -0.2659 Tw ( ) Tj ET 120.75 585.75 m 120.75 585 l 121.5 585 l 121.5 584.25 l 120.75 584.25 l 120.75 584.25 l 120 584.25 l 120 585.75 l 120.75 585.75 l 120.75 583.5 m 119.25 577.5 l 118.5 576.75 l 118.5 576 l 117.75 576 l 117.75 575.25 l 117 575.25 l 116.25 574.5 l 116.25 575.25 l 115.5 575.25 l 115.5 576 l 116.25 576 l 116.25 575.25 l 117.75 575.25 l 117.75 576.75 l 119.25 581.25 l 119.25 582.75 l 118.5 582.75 l 120.75 583.5 l h f* 0.1216 0.102 0.0902 rg 91.5 728.25 m 91.5 726.75 l 90 726.75 l 90 728.25 l h f 94.5 728.25 m 94.5 726.75 l 93 726.75 l 93 728.25 l h f 97.5 728.25 m 96 728.25 l 96 726.75 l 97.5 726.75 l h f 100.5 728.25 m 100.5 726.75 l 99 726.75 l 99 728.25 l h f 103.5 728.25 m 102 728.25 l 102 726.75 l 103.5 726.75 l h f 106.5 728.25 m 106.5 726.75 l 105 726.75 l 105 728.25 l h f 109.5 728.25 m 108 728.25 l 108 726.75 l 109.5 726.75 l h f 112.5 727.5 m 112.5 726.75 l 111 726.75 l 111 728.25 l h f 115.5 727.5 m 114 727.5 l 114 726 l 115.5 726 l h f 118.5 727.5 m 118.5 726 l 117 726 l 117 727.5 l h f 121.5 727.5 m 120 727.5 l 120 726 l 121.5 726 l h f 124.5 726.75 m 124.5 725.25 l 123 726 l 123 727.5 l h f 127.5 726.75 m 126.75 725.25 l 125.25 725.25 l 126 726.75 l h f 130.5 726.75 m 129.75 725.25 l 128.25 725.25 l 129 726.75 l h f 133.5 726 m 132.75 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